VILNIUS, Lithuania—A week after Russia’s leading opposition figure Alexei Navalny died in an Arctic prison colony in February, his wife, Yulia Navalnaya, met with his grieving aides to ask: What next?

       It was a pressing question not just for an organization that, despite Navalny’s star power, had struggled to dent President Vladimir Putin’s regime, but for the Russian opposition as a whole.

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