Koalas — Australia’s iconic marsupials — have been petted by princes, pop stars, presidents and a pope. But one of the country’s oldest koala sanctuaries has stopped offering cuddles with the furry animals in response to strong visitor feedback.

       The move by Brisbane’s Lone Pine Koala Sanctuary highlights a growing debate about whether the welfare risks to animals of “selfie-style” tourist encounters — those centered on photo opportunities — outweigh the benefits of encouraging conservation.