KRAMATORSK, Ukraine —

       Thunderous blasts echoed in the capital, Kyiv, and in Ukraine’s eastern battle zone on Friday, a day after Russia’s most important warship in the Black Sea sank while being towed to port.

       Russia’s defense ministry threatened the stepped-up missile attacks after its flagship missile cruiser Moskva went down Thursday following what Ukraine said was a missile strike and what Moscow claimed was an accidental fire aboard.

       The capital had enjoyed a period of relative calm after Russian forces broke off an offensive in Ukraine’s north at the beginning of this month, although departing Russian troops left behind evidence of mass atrocities committed during a month-long occupation of some of Kyiv’s satellite towns and suburbs.

       Powerful explosions were also heard Friday in Kramatorsk, in Ukraine’s east, Kherson in the south, and in the northeastern city of Kharkiv, in advance of an expected major Russian onslaught in the industrial eastern heartland known as the Donbas.

       Ukraine’s armed forces, meanwhile, claimed to have repelled Russian attacks on two towns, Popasna and Rubizhne, near the embattled southern port city of Mariupol.

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       Ukraine said outgunned defenders were still holding on in Mariupol, which Moscow is determined to capture in order to create a land bridge between Russian-controlled territory and the Crimean Peninsula, seized by Russia in 2014. But analysts predicted the city’s fall could be near.

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       Ukrainian President Volodymyr Zelensky, in his overnight address to the nation, praised Ukrainian defenders’ perseverance and courage as the war entered its eighth week – an invasion the Ukrainian leader described as “absurd” and “suicidal.”

       He also mocked the loss of the Moskva. He did not refer to the cruiser by name, but declared that Ukrainian forces had shown by their actions that Russian warships go “to the bottom only.”

       The flagship’s loss will likely force changes in how the Russian navy operates in the Black Sea, a major staging ground for Moscow’s expected offensive in Ukraine’s east and its southern seacoasts, analysts said. In an assessment on Friday, British military intelligence said the Moskva had served a crucial role as command vessel and air defense node.

       Combined with Ukraine’s attack in late March on the landing ship Saratov, Russia “has now suffered damage to two key naval assets” since the Feb. 24 invasion, the British assessment said, adding that those two episodes “will likely lead Russia to review its maritime posture in the Black Sea.”

       However, a Washington think tank, the Institute for the Study of War, said in its latest analysis that while the Moskva’s loss will reduce Russia’s ability to conduct cruise missile strikes, it is “unlikely to deal a decisive blow to Russian operations as a whole.”

       Other Russian vessels have swiftly moved away from the southern Ukraine coastline, at least for now, Western military officials said. But the Pentagon said that on land, dozens of Russian battalion tactical groups are in place for the expected eastern offensive.

       Zelensky, in his overnight address, harkened back to the start of the invasion, which he said Russia expected to make short work of subduing Ukraine.

       Marking 50 days under attack, he called the country’s defense “an achievement of millions of Ukrainians, of everyone who on Feb. 24 made the most important decision of their life — to fight.”

       In vowing renewed strikes on the capital, Moscow did not cite the sinking of the Moskva – although it did acknowledge the warship’s loss -- but instead said it was responding to attacks on Russian territory.

       “The number and scale of missile strikes against targets in Kyiv will increase in response to the Kyiv nationalist regime committing any acts of a terrorist nature or sabotage on Russian territory,” Defense Ministry spokesman Igor Konashenkov said, according to the RIA news agency.

       Russian authorities on Thursday had accused Ukraine of a cross-border helicopter attack on residential buildings in the village of Klimovo, in Russia’s Bryansk region.

       Speaking on the BBC, former CIA director David Petraeus described Russia’s admission that the ship sank as a “rare moment of truth,” but said facts surrounding the vessel’s loss would have “come out” eventually.

       Bulos reported from Kramatorsk and King from Warsaw, Poland.